A wasp is any insect of the order Hymenoptera and suborder Apocrita that is neither a bee nor an ant. This means that wasps are paraphyletic with respect to bees and ants, and that all three groups are descended from a common ancestor; the Apocrita form a clade.
The most commonly known wasps such as yellow jackets and hornets are in the family Vespidae and are eusocial, living together in a nest with an egg-laying queen and non-reproducing workers. Eusociality is favoured by the unusual haplodiploid system of sex determination in Hymenoptera, as it makes sisters exceptionally closely related to each other. However, the majority of wasp species are solitary, with each adult female living and breeding independently. Many of the solitary wasps are parasitoidal, meaning that they raise their young by laying eggs on or in the larvae of other insects. The wasp larvae eat the host larvae, eventually killing them. Solitary wasps parasitize almost every pest insect, making wasps valuable in horticulture for biological pest control of species such as whitefly in tomatoes and other crops.
Wasps first appeared in the fossil record in the Jurassic, and diversified into many surviving superfamilies by the Cretaceous. They are a successful and diverse group of insects with tens of thousands of described species; wasps have spread to all parts of the world except for the polar regions. The largest social wasp is the Asian giant hornet, at up to 5 centimetres (2.0 in) in length; among the largest solitary wasps is the giant scoliid of Indonesia, Megascolia procer. The smallest wasps are solitary chalcid wasps in the family Trichogrammatidae, some of which are just 0.2 millimetres (0.008 in) in length.
Wasps play many ecological roles. Some are predators, whether to feed themselves or to provision their nests. Many, notably the cuckoo wasps, are kleptoparasites, laying eggs in the nests of other wasps. With their powerful stings and conspicuous warning coloration, often in black and yellow, wasps are frequent models for Batesian mimicry by non-stinging insects, and are themselves involved in mutually beneficial Müllerian mimicry of other distasteful insects including bees and other wasps. Wasps have appeared in literature from Classical times, as the eponymous chorus of old men in Aristophanes' 422 BC comedy Σφῆκες (Sphēkes), The Wasps, and in science fiction from H. G. Wells's 1904 novel The Food of the Gods and How It Came to Earth, featuring giant wasps with three-inch-long stings. The name "Wasp" has been used for many warships and other military equipment.
Relationship with humans
As pests
Social wasps are considered pests when they become excessively common, or nest close to buildings. People are most often stung in late summer, when wasp colonies stop breeding new workers; the existing workers search for sugary foods and are more likely to come into contact with humans; if people then respond aggressively, the wasps sting. Wasp nests made in or near houses, such as in roof spaces, can present a danger as the wasps may sting if people come close to them. Stings are usually painful rather than dangerous, but in rare cases people may suffer life-threatening anaphylactic shock.
In horticulture
Further information: biological pest control
Some species of parasitic wasp, especially in the Trichogrammatidae, are exploited commercially to provide biological control of insect pests. For example, in Brazil, farmers control sugarcane borers with the parasitic wasp Trichogramma galloi. One of the first species to be used was Encarsia formosa, a parasitoid of a range of species of whitefly. It entered commercial use in the 1920s in Europe, was overtaken by chemical pesticides in the 1940s, and again received interest from the 1970s. Encarsia is used especially in greenhouses to control whitefly pests of tomato and cucumber, and to a lesser extent of aubergine (eggplant), flowers such as marigold, and strawberry. Several species of parasitic wasp are natural predators of aphids and can help to control them. For instance, Aphidius matricariae is used to control the peach-potato aphid.
In sport
Wasps RFC is an English professional rugby union team originally based in London but now playing in Coventry; the name dates from 1867 at a time when names of insects were fashionable for clubs. The club's first kit is black with yellow stripes. The club has an amateur side called Wasps FC.
Among the other clubs bearing the name are a basketball club in Wantirna, Australia, and Alloa Athletic F.C., a football club in Scotland.
In fashion
Wasps have been modelled in jewellery since at least the nineteenth century, when diamond and emerald wasp brooches were made in gold and silver settings. A fashion for wasp waisted female silhouettes with sharply cinched waistlines emphasizing the wearer's hips and bust arose repeatedly in the nineteenth and twentieth centuries.
In literature
The Ancient Greek playwright Aristophanes wrote the comedy play Σφῆκες (Sphēkes), The Wasps, first put on in 422 BC. The "wasps" are the chorus of old jurors.
It flew, he is convinced, within a yard of him, struck the ground, rose again, came down again perhaps thirty yards away, and rolled over with its body wriggling and its sting stabbing out and back in its last agony. He emptied both barrels into it before he ventured to go near. When he came to measure the thing, he found it was twenty-seven and a half inches across its open wings, and its sting was three inches long. ... The day after, a cyclist riding, feet up, down the hill between Sevenoaks and Tonbridge, very narrowly missed running over a second of these giants that was crawling across the roadway.
Wasp is a 1957 science fiction book by the English writer Eric Frank Russell; it is generally considered Russell's best novel.
Parasitoidal wasps played an indirect role in the nineteenth-century evolution debate. The Ichneumonidae contributed to Charles Darwin's doubts about the nature and existence of a well-meaning and all-powerful Creator. In an 1860 letter to the American naturalist Asa Gray, Darwin wrote:
I own that I cannot see as plainly as others do, and as I should wish to do, evidence of design and beneficence on all sides of us. There seems to me too much misery in the world. I cannot persuade myself that a beneficent and omnipotent God would have designedly created the Ichneumonidae with the express intention of their feeding within the living bodies of caterpillars, or that a cat should play with mice.
In military equipment
With its powerful sting and familiar appearance, the wasp has given its name to many ships, aircraft and military vehicles. Nine ships and one shore establishment of the Royal Navy have been named HMS Wasp, the first an 8-gun sloop launched in 1749. Eleven ships of the United States Navy have similarly borne the name USS Wasp, the first a merchant schooner acquired by the Continental Navy in 1775. In the Second World War, a German self-propelled howitzer was named Wespe, while the British developed the Wasp flamethrower from the Bren Gun Carrier. In aerospace, the Westland Wasp was a military helicopter developed in England and used by the Royal Navy and other navies; it first flew in 1958. The AeroVironment Wasp III is a Miniature UAV developed for United States Air Force special operations.
In astronomy
WASP is the acronym for Wide Angle Search for Planets; this has in turn led to 'WASP' becoming part of the designations of the many planets it has discovered.
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